Yonas Molar Site
Basic information
Sample name: Yonas Molar Site

Sample aka: ARA-VP-1

Reference: T. D. White, S. H. Ambrose, G. Suwa, D. F. Su, D. DeGusta, R. L. Bernor, J.-R. Boisserie, M. Brunet, E. Delson, S. Frost, N. Garcia, I. X. Giaourtsakis, Y. Haile-Selassie, F. C. Howell, T. Lehmann, A. Likius, C. Pehlevan, H. Saegusa, G. Semprebon, M. Teaford, and E. Vrba. 2009. Macrovertebrate paleontology and the Pliocene habitat of Ardipithecus ramidus. Science 326:87-93 [ER 4407]
Geography
Country: Ethiopia



Coordinate: 10.477° N, 40.446° E
Coordinate basis: estimated from map

Scale: quarry

Formation: Sagantole

Time interval: Early Pliocene

Max Ma: 4.419

Min Ma: 4.416

Age basis: Ar-Ar

Geography comments: the site name is given by White et al. (2009)
from the Lower Aramis Member of the Sagantole Formation in "the Central Awash Complex (CAC) of the Middle Awash study area" (WoldeGabriel et al. 2009)
the exact location is shown in Fig. 1 of WoldeGabriel et al. (2009
"The Aramis Member directly overlies the Gŕala (“Camel”) Tuff Complex (GATC), which has a 40Ar/39Ar age of 4.419 ± 0.068 Ma" and is "overlain by the Daam Aatu (“Baboon”) Basaltic Tuff (DABT), which has a 40Ar/39Ar age of 4.416 ± 0.031 Ma" (WoldeGabriel et al. 2009)

Environment
Lithology: claystone

Taphonomic context: overbank deposit,bird accumulation

Habitat comments: the member consists of "silt, clay, and sand of variable thickness and induration, deposited on a floodplain" (WoldeGabriel et al. 2009) and the fossils are specically from "alluvial silty clay" according to White et al. (2009)
there is no evidence of "hydraulic transport" but is marking of bone by mammalian carnivores, crocodiles, rodents, and insects (Louchart et al. 2009)

Methods
Life forms: carnivores,primates,ungulates,other large mammals

Sampling methods: quarry,screenwash

Sample size: 1342 specimens

Years: 1992 - 1997

Sampling comments: quarried, screenwashed, and surface crawled
collection years are mentioned in Fig. 2 of White et al. (2009) and in the text
indeterminate bovids identified to tribe are also enumerated
small mammals and bird bones are present according to Louchart et al. (2009) and pollen and millipedes are present according to WoldeGabriel et al. (2009), who also suggest that "insect larvae, dung beetle broodballs and nests, diverse gastropods" are present
"wood fragments, seeds" were collected according to White et al. (2009)

Metadata
Sample number: 4686

Contributor: John Alroy

Enterer: John Alroy

Modifier no: John Alroy

Created: 2025-08-30 06:40:02

Modified: 2025-08-30 06:46:21

Abundance distribution
32 species
6 singletons
total count 1342
geometric series index: 46.8
Fisher's α: 5.890
geometric series k: 0.8141
Hurlbert's PIE: 0.7581
Shannon's H: 2.0098
Good's u: 0.9955
Each square represents a species. Square sizes are proportional to counts.
Register
Hippopotamidae indet.15
Cainochoerus sp.4
said to be new
Kolpochoerus deheinzelini11
Nyanzachoerus jaegeri = †Notochoerus jaegeri6
Nyanzachoerus kanamensis35
Neotragini indet.12
Aepyceros sp.51
cf.
Damalops sp.6
Gazella sp.1
questionably
Simatherium sp.4
cf.
Ugandax cf. gautieri3
also 2 Ugandax sp.
Tragelaphus cf. moroitu588
referred to in the table as "Tragelaphus kyloae" (a nomen nudum)
Praedamalis sp.1
Kobus sp.1
"Kobus preoricornis" (an apparent nomen nudum)
Giraffa sp.8
Sivatherium sp.9
Hyaenidae indet.42
Enhydriodon sp.1
Agriotherium sp.2
Dinofelis sp.1
questionably
Felis "small"6
Felis "medium"2
also 2 "Felis indet."
Ceratotherium sp.5
said to be new
Diceros sp.1
Eurygnathohippus sp.45
Pliopapio alemui164
Kuseracolobus aramisi225
Ardipithecus ramidus54
Anancus sp.3
Deinotherium sp.2
Elephantidae indet.10
Orycteropus sp.24