Trlica (Bed 5)
Basic information
Sample name: Trlica (Bed 5)

Reference: A. K. Agadzhanyan, I. A. Vislobokova, M. V. Shunkov, and V. A. Ulyanov. 2017. Pleistocene mammal fauna of the Trlica locality, Montenegro. Fossil Imprint 73(1-2):93-114 [ER 3942]
Geography
Country: Montenegro



Coordinate: 43° 20' 38" N, 19° 23' 0" E
Coordinate basis: stated in text

Time interval: Middle Pleistocene

Unit number: 5

Unit order: above to below

Max Ma: 0.79

Min Ma: 0.676

Age basis: zone

Geography comments: "2.5 km southeast of the town of Pljevlja and 1.6 km northeast of the mouth of the Cehotina River canyon"
"not older than 1 Ma" and from "the first half of he Middle Pleistocene... starting from MIS 19" based on biochronology: shown on a chart as within the range of MIS 19 to MIS 17 and roughly 0.7 to 0.8 Ma, consistent with the boundaries of these two stages

Environment
Lithology: siliciclastic (mixed)

Taphonomic context: carnivore accumulation,cave,rodent accumulation

Habitat comments: a "closed karstic cavity filled with loose deposits" and this bed is specifically "Ocherous-yellow heavy loam and yellow intermediate loam, with a brown tint... The middle part of the bed displaces traces of activity by large digging animals. The lower part of the bed contains strongly weathered pebbles... Teeth and bones of large mammals are regularly scattered throughout... small mammals... are rather scarce. The bed is up to 1.0-1.3 m in total thickness" and was "formed by diluvial-proluvial processes"
the coprolites are "of hyenas... the cave was periodically inhabited and occupied at least by hyenas and porcupines"
human artefacts are not mentioned and presumably absent

Methods
Life forms: carnivores,rodents,ungulates

Sampling methods: quarry,screenwash

Sample size: 109 specimens

Years: 2010 - 2014

Sampling comments: "all sediments taken out of the cave were washed in sieves with 1 x 1 mm mesh"

Metadata
Sample number: 4346

Contributor: John Alroy

Enterer: John Alroy

Created: 2023-08-18 15:36:49

Modified: 2023-08-18 05:36:49

Abundance distribution
21 species
5 singletons
total count 109
geometric series index: 38.3
Fisher's α: 7.738
geometric series k: 0.8679
Hurlbert's PIE: 0.9043
Shannon's H: 2.6175
Good's u: 0.9551
Each square represents a species. Square sizes are proportional to counts.
Register
Ursus deningeri1
plus 1 Carnivora and 1 Ursus
Canidae indet.2
one of which is possibly Vulpes or Nyctereutes
Elephantidae indet.3
Stephanorhinus hundsheimensis13
plus 1 Rhinocerotidae
Equus cf. suessenbornensis2
plus 2 Equus
Equus sp.2
small, not the same: plus 3 Equidae
Praemegaceros cf. verticornis2
plus 4 Praemegaceros
Cervus elaphus acoronatus17142 kg browser-grazer
"Cervus cf. acoronatus": plus 3 Cervus
Capreolus sp.1
questionably referred
Bison cf. schoetensacki13
plus 6 Bison
Soergelia sp.6
plus 7 Ovibovini and 1 Bovidae
Glis sackdillingensis17
Cricetus sp.6
Cricetulus sp.2
Mimomys ex gr. intermedius7
Mimomys cf. pusillus4
plus 6 Mimomys
Microtus (Terricola) hintoni6
"Terricola hintoni"
Microtus (Terricola) arvaloides2
"Terricola arvaloides": plus 1 Terricola and 14 Microtini
Lagurodon cf. arankae1
Lagurus ex gr. transiens1
plus 9 Lagurus
Hystrix sp.1
plus 98 Rodentia